Apple Facts & Health Benefits: Everything About Apples

If there’s an iconic fruit, it’s the apple. From the Big Apple to “American as apple pie” to your sweetheart being the “apple of your eye,” this delicious fruit has captured our imaginations like no other. It’s both revered for its health benefits (keeping the doctor away) and reviled for its contribution to original sin (although the fruit in the Garden of Eden is no longer thought to have been an apple.)

Apples fill not just our thoughts and religious traditions, but our shopping carts as well. The apple and its comical buddy, the banana, are the most consumed fruits in the United States. (Worldwide, the apple is hugely popular too. Although it loses ground to the tomato — if we’re going to call tomatoes fruits, that is!)

And no wonder apples are popular. They’re not only delicious but convenient as well. They’re portable, pre-wrapped, long-lasting, relatively inexpensive, and widely available. And, as we’ve seen, they’ve been imbued by our collective imagination with both wholesome and health-giving properties. But there are some concerns, including their high sugar content, pesticide exposures, and more and more genetically modified varieties are coming out. So all in all, how good for you are apples? Do their benefits outweigh any downsides? Let’s examine the apple facts.

Apple Facts

According to DNA analysis, apples originated in the mountains of Kazakhstan, in Central Asia. Apples are members of Rosaceae, the Rose family, along with pears, cherries, strawberries, raspberries, plums, and peaches. Apple trees reproduce sexually, meaning that both a male and female tree are required to produce a viable seed. And here’s the crazy thing. Unlike most plants, baby apple trees can produce fruit completely differently from either parent — an evolutionary strategy known as “extreme heterozygosity.” Pretty ironic, considering the meaning of the phrase, “The apple doesn’t fall far from the tree!”

These qualities of huge variance and utter unpredictability in the propagation of apples are what made Johnny Appleseed so important to the history of the apple. By planting seeds everywhere he went in Pennsylvania, Ohio, West Virginia, Ontario, and Illinois, he provided the seed stock for many of the varieties of apples we enjoy today. But because farmers can’t count on new trees from seed to produce edible and marketable fruit, their only guarantee of reproducibility is grafting, which is how the modern apples we’re used to eating come about. Grafting apples is a process by which buds are carefully removed from the current year’s growth and inserted into the bark of trees used to grow the following year’s apples. A well-cared-for apple tree can live from 100 to 200 years.

Types of Apples

When you go to the grocery store, chances are you’re familiar with the most common types of apples to choose from — for instance, Golden, Pippin, Granny Smith, and Fuji. And don’t forget Red Delicious, the most well-known of all, which reigned as the most popular apple in the United States for decades until 2018 when it was dethroned by the Gala apple. But did you know that worldwide there are over 7,500 cultivated apple types?

The most popular cultivated apple types include:

Red Delicious
Granny Smith
Gala
Fuji
Honeycrisp
Golden Delicious
McIntosh
Pink Lady/Cripps Pink
Braeburn
Idared
Cosmic Crisp

Rare Apples

Although most apples you see in the grocery store are red, pink, yellow, or green with a whitish interior, rarer apples may be black or purple on the outside, or have red or pink flesh. Some even taste like roses, lemon, or honey. What’s even more interesting is that the sweeter apples most of us are used to weren’t the original apples. Today’s commercial apples were likely only introduced within the last two hundred years.

Rarer apples can be expensive as they’re not often commercially grown. The Tibetan Black Diamond apple — which is a dark purple apple that grows in a very remote part of Tibet and resembles something from a fairy tale — can cost up to $20/apple. Most of these rare black apples never make it to the consumer marketplace, partially because apple farmers are uncertain how well they would sell internationally, and because they can take five to eight years to even bear fruit.

If you’re not planning to travel to Tibet anytime soon, a similar dark purple apple grows in Arkansas and is called the Arkansas Black apple. Still, this one is too tart to eat when first picked. And it requires long-term storage to develop its flavor, which prevents it from being widely available to consumers.

Apple Nutrition

Nearly all apple varieties are a good source of important nutrients, especially fiber, vitamin C, and potassium, as well as various antioxidants and phytochemicals — such as quercetin, catechin, and chlorogenic acid.

Much of the fiber in apples is called pectin, which is a mixture of soluble and insoluble fibers. Soluble fiber helps feed good gut bacteria and lower high cholesterol. While insoluble fiber helps keep your intestines clean and healthy.

Apples contain simple natural sugars like glucose, fructose, and sucrose, but they have a low glycemic index (GI) — around 36 — which means they don’t spike your blood sugar much after you eat them. While some people avoid apples because they are so sweet, their high fiber and polyphenol content actually help prevent glucose spikes by slowing down how fast your body breaks down carbohydrates.

One medium (3-inch diameter) apple offers the following nutritional profile:

Calories: 95
Protein: 0.5 grams
Total Fat: 0.3 grams
Total Carbohydrate: 25 grams
Fiber: 4.5 grams
Vitamin C: 14% of the Daily Value (DV)
Potassium: 6% of the DV
Vitamin K: 5% of the DV
Vitamin B6: 4% of the DV

Keep in mind that there are many nutritional components in the apple peel, so it’s best to eat them washed and unpeeled.

Health Benefits of Apples

In addition to being packed with nutrients, apples offer a number of health benefits. Discover some of the benefits with the most research support below.

1. May have anticancer benefits.

Some of the phytonutrients in apples appear to have protective effects when it comes to cancer risk. A 2005 study published in the Annals of Oncology reviewed studies done between 1991 and 2002 in Italy, consistently finding that people who consumed at least one apple per day were at a 20% and 18% lower risk of colorectal and breast cancers, respectively. Other research suggests that polyphenolic compounds in apples have antioxidant, anti-inflammatory, anti-proliferative, and apoptosis-inducing characteristics when it comes to cancer.

2. May lower your risk for cardiovascular disease.

Human and animal studies have found that regularly eating apples can protect heart health. A 2008 study published in Molecular Nutrition & Food Research fed some very unfortunate hamsters a high cholesterol diet for 12 weeks. When these hamsters also received apples and apple juice, they experienced a reduction in cholesterol levels and a 48% reduction in arterial plaque buildup compared to a control group. The researchers attributed this effect partially to the antioxidant boost the hamsters received from the fruit.

But what about humans? A Finnish study from 1998 found that men and women who ate more than 54 grams of apples daily (about half a decent-sized apple) lowered their risk of dying from a heart attack by 19% and 43%, respectively. And more recently, a 2020 study published in the American Journal of Clinical Nutrition fed healthy, mildly hypercholesterolemic adults either two apples per day or a sugar- and energy-matched apple control beverage for eight weeks each. Researchers measured heart health biomarkers before and after each treatment. They found that eating two apples per day reduced total cholesterol, LDL “bad” cholesterol, and triglycerides compared to the control group.

3. May support healthy weight loss.

Apples are high in fiber and low in calories, which are two characteristics that make them an ideal weight-loss food. Their fiber content can contribute to satiety, preventing overeating during the day, and reducing overall caloric intake. In a 2003 study, overweight women between ages 30-50 years with high blood cholesterol levels were randomized to eat supplements of either apples, pears, or oat cookies for 12 weeks. The group who consumed one and a half apples (300 grams) per day lost three pounds during the course of the study. (To be fair, the pear group also lost weight, but hey, this article is about apples. Pears, wait your turn.) A 2018 review published in the Journal of the American College of Nutrition looked at current research around apple components and weight loss, finding that there is a consistent correlation between apple consumption and weight loss.

4. May improve lung health and asthma symptoms.

Consuming fruits and vegetables, in general, has been found to have a positive influence on lung health. In 2007, an in-depth analysis of the link between produce consumption and lung cancer, based on data collected from a prospective cohort of 478,590 participants from 10 European countries, was published. A significant inverse association occurred between daily intake of apples and pears and lung cancer incidence. In other words, the more apples (and pears — there they go trying to creep in again!) people ate, the less likely they were to get lung cancer.

5. May lower the risk of dementia and protect brain health.

A 2005 study in the Journal of Alzheimer’s Disease found that mice fed apple juice concentrate after being fed a pro-oxidant diet (deficient in vitamin E and folate, but high in iron) had significantly improved cognition and reduced pro-oxidative status compared to the control group. The amount fed to the mice was the equivalent of two to three, eight-ounce glasses of apple juice per day for humans. Other research suggests that apple juice concentrate may intervene in mechanisms that promote production of amyloid β peptide, a hallmark of Alzheimer’s.

6. May support healthy blood sugar control.

It’s counterintuitive, considering how sweet apples are, but a growing body of research suggests that eating apples may help lower high blood sugar and protect against type 2 diabetes. Furthermore, the fiber and polyphenols in apples are thought to slow carbohydrate digestion, preventing dramatic spikes in blood sugar after eating. In a 2005 study published in the Journal of the American College of Nutrition, researchers examined the association of dietary flavonoid intake and type 2 diabetes among 38,000 women, finding that eating one or more apples per day was linked to a 28% lower risk of developing type 2 diabetes.

7. May improve bone health.

Research shows that eating fruits and vegetables improves bone mineral density and other markers of bone strength. In a 2004 clinical trial, 15 healthy female adults ate a 500-calorie, macronutrient-matched test meal on three different occasions, which either included fresh peeled apples, unsweetened applesauce, or candy. Urine samples were taken after the meals, finding that eating fresh apples and applesauce each reduced net acid excretion from the body, and slowed calcium loss.

8. May help protect your gastrointestinal system.

Apples may not only protect your gut via their fiber composition; they may actually protect it from damage caused by medications. For instance, a 2005 study published in the journal Gut found that apple extract helped prevent damage to the gut known to be a risk factor from using non-steroidal anti-inflammatory drugs (NSAIDs). This attribution came from the polyphenols catechin and chlorogenic acid, which help prevent oxidative damage to cells in the body. Preliminary test-tube studies have also indicated that apple polyphenols may protect against gastric ulcers.

9. May stimulate hair growth.

It turns out that there may be some truth to the touted growth-promoting effects of certain apple-derived hair products. A 2002 study published in the British Journal of Dermatology observed the effects of apple-derived procyanidin B-2 on the growth of hair in rats, concluding that it has a hair growth-promoting mechanism and plays a role in the hair cycle progression. Ironic, considering how smooth and shiny apples are!

5 Common Apple Criticisms and Potential Health Risks

While apples offer undeniable health benefits, they’ve also received criticism for risks that may come with eating them. Are these concerns warranted or not? Let’s examine some of them and separate apple facts from fiction.

1. Apples are high in cyanide.

While it’s true that apple seeds contain amygdalin — a cyanide and sugar compound that degrades into hydrogen cyanide when metabolized — the human body can detoxify this in small doses. I don’t know about you, but most people don’t even eat the apple seeds — at least, not on purpose. And the number of apple seeds it would take for the cyanide content to become dangerous is so large that even apple seed lovers are unlikely to be at risk for issues. Plus, you’d have to chew them really well to release the amygdalin into your bloodstream. Creating a lethal apple seed concoction would probably require grinding the seeds from hundreds of apples and consuming them all at once. (Don’t do that!) The good news: apple flesh and peel don’t contain any cyanide.

2. Apples are high in sugar.

Like other fruits and vegetables, apples do contain their share of natural sugars. Sugars and carbohydrates often get a bad rap for people with prediabetes or diabetes. But apples don’t actually contribute to this disease. The average apple contains 25 grams of carbohydrates, 19 grams of which are sugars. But apples are low on the glycemic index because of their fiber and polyphenolic content. Note, however, that this is referring to whole apples. Juiced apples are another matter as apple juice has had most — if not all — of its natural fiber filtered out and discarded, leaving you with a sugar-rich liquid that will likely affect your blood sugar differently.

3. Apples are often genetically modified.

One of the most common genetic modifications done to apples is to prevent them from browning. Arctic apples, for instance, have this modification. While it doesn’t improve their flavor or nutritional value, the flesh of these apples won’t turn brown when sliced and exposed to oxygen like other apples will. This is appealing to food service providers and some restaurants because they can leave sliced apples out for longer without them turning brown.

Arctic apples could lead to some degree of reduction in food waste. But they lead to no net improvement in flavor, nutritional value, or yield. And they don’t help reduce water or pesticide consumption, either. And GMO apples that don’t brown could lead to people unknowingly consuming apples that could be weeks or even months old or more. Are there risks? We don’t know. And without proper labeling, we likely never will. Purchase organic or non-GMO labeled apples whenever possible to reduce pesticide exposure and opt-out of the GMO experiment.

4. Apples contain high amounts of pesticides.

Every year, the Environmental Working Group (EWG) releases their shopper’s guide to pesticides in produce, called the Dirty Dozen™. And every year, apples make the list. According to a 2019 article from EWG, apples contain at least four pesticide residues on average, including some at high concentrations. The use of some pesticides occurs after apples are harvested, like diphenylamine, which doesn’t have enough long-term safety data available at this time. That’s why I suggest buying organic apples whenever possible. And to help remove any pesticide residues, soak them in a mixture of water and baking soda, or at the very least scrub them thoroughly with a produce brush before eating.

5. Apples are a source of allergens.

Although rare, there is such a thing as an apple allergy. Specifically, it’s an allergy to the protein in apples. What may be more common, however, is a reaction called oral allergy syndrome (OAS), which can appear suddenly after eating certain fresh produce like apples. Symptoms typically include itchy lips and tongue. But OAS isn’t actually an allergy to apples themselves; it’s an overly-protective bodily response to pollen. In a sense, your immune system mistakes the apple for pollen that you’re allergic to. OAS is more common among people with seasonal allergies or hay fever who react to pollen in the environment. Cooking produce can remove the allergen, but an OAS response typically only lasts 15 minutes and isn’t usually severe.

How to Choose Apples

Like other fresh produce, proper storage will help preserve apples in your home for as long as possible. Furthermore, knowing what to look for at the grocery store or farmers market will help you choose the best quality apples.

When purchasing apples, look for any damage or discoloration on the skin. When you feel apples in your hand, avoid any that have soft or mushy spots. And after you get your apples home, eat the ones that have any damage first as these will start to go bad faster the longer they sit. Remember what they say about “one bad apple”!

Tips for Apple Storage

At home, consider how long you’ll be storing your apples before eating them. Apples with thinner skins don’t last as long. Furthermore, apples give off a compound called ethylene gas, which will speed up the ripening of other produce. Hence, apples should usually be stored away from other produce, in your fridge or on your countertop. The ideal temperature for longest storage is 30-35°F (-1-1.5°C) with 90-95% relative humidity. If storing a small amount in the fridge, place the apples in a crisper drawer in a plastic bag with holes, or cover them with a damp paper towel. Putting apples in an airtight container or sealed bag right away will release moisture and make them rot quicker.

If you have a lot of apples to store, a cool, dark place with a little bit of humidity, like a basement or garage, may be a good choice. For very long-term storage, the most optimal method is to individually wrap apples in paper with their stem still intact. The paper helps keep the apples separate to prevent them from ripening each other. Wrapped apples should be put in a crate, ideally in a single layer, or in trays. Be sure to wash the apples well before eating them to remove any paper residue.

The Best Ways to Eat Apples

The best way to enjoy apples depends on the specific variety. Some types of apples are firmer, have thinner skin, or break down more easily than others.

Here are some suggested best uses for some of the most popular types of apples (all of which are also good eaten straight):

McIntosh: These break down easily and are a good choice for homemade applesauce.
Fuji: This type is sturdy and can withstand baking and roasting, making it ideal for strudel or apple crisp.
Red Delicious: These are sensitive to heat and are best enjoyed fresh and crisp, like sliced into salads or crunchy sandwiches.
Gala: This type is also heat-sensitive and works best raw or cut and tossed into fruit salads.
Crispin: This variety is versatile and works well baked, in apple pies, or made into applesauce.
Braeburn: The sweetness of this type mellows when heated. But the Braeburn apple keeps its shape. Braeburns work well in both sweet and savory dishes.
Honeycrisp: This apple remains sweet even when cooked, so it does well in dishes like crisps and pies, but also slaws and sandwiches.
Granny Smith: This tart, high-acidity variety is tasty raw or cooked into pies, tarts, pancakes, soups, or stuffing.

Enjoy Eating Apples

Regardless of the type, apples are one of the most versatile fruits. And there are many ways to enjoy them. Try adding chopped apples to cold cereal or hot oatmeal, in plant-based yogurt with other healthy toppings, on pancakes or waffles, sliced and stacked onto peanut butter sandwiches, or simply sliced and served with nut butter or a homemade chickpea-based “cookie dough” dip. You can boil apples, mash them into homemade cinnamon applesauce, grate them into coleslaw, grill, stuff, and bake them, or dice and mix them into cold salads. Apples are a healthy, delicious addition to just about any dish or snack. And, of course, you can also eat them raw, straight away, without any preparation (other than rinsing in clean water).

Healthy Apple Recipes

Let’s take a moment to appreciate the versatility of apples. They can add a little sweetness and crunch to salads, add texture to and develop flavor in salad dressings, or you can bake them into a delicious dessert. It’s time to step away from the traditional apple and peanut butter (though I think it’s one of the most delicious snacks on the planet!) and experiment with all of the ways to use apples in recipes. To get you started, we’ve created three for you to make and enjoy.

1. Kale Pomegranate Salad with Chopped Walnuts and Apples

Apples can lend a subtle sweetness to savory dishes like this Kale Pomegranate Salad, where it pairs perfectly with the acidic dressing and earthy kale. The vitamin C in the apples can also increase the iron absorption from the kale. And you can enjoy this salad year-round, from a refreshing meal on a summer evening to a colorful dish for a winter holiday party.

2. Magical Applesauce Dressing

Who knew that apples could help emulsify a dressing? When you change the consistency from whole apple to applesauce, apples make for a perfect dressing ingredient by adding texture and natural sweetness.

3. Apple Crumble

Have a sweet tooth but don’t want the overly processed sugar? This Apple Crumble might help to satisfy it naturally. Braising the apples helps to bring out their natural sugar. And adding a bit of date paste adds more natural sweetness, plus fiber and nutrition.

Fact: An Apple a Day is a Good Start

Apples are a fantastic addition to a healthy, whole foods, plant-based diet. It’s no wonder they’re so well-loved, considering they have many benefits and offer properties that can help prevent chronic disease. While there are some “bad apples” out there — like GMOs and pesticide-laden crops — the real apple facts show there’s definitely some truth to the adage of “an apple a day keeps the doctor away.”

Tell us in the comments:

What’s your favorite variety of apple, and why?
What apple-based recipes do you like to make at home?
How do you store apples?

Feature image: iStock.com/cometary

Apple Facts You Need To Know About: Benefits, Concerns, & the Best Ways to Enjoy Them

Ocean Robbins